Introduction
Animals, including humans, are born with in-built instincts to perform adaptive behaviors. These instincts include many reflexes and relatively straightforward behaviors, such as food-seeking behavior. But, as animal behavior gets more complex, there are in-build instincts which are correspondingly more complex (e.g., food storing behaviors). Thus, personality in humans is considered, from an evolutionary perspective, not be qualitatively different in origin than the drive in dogs, for example, to bury their bones, or squirrels to store acorns. It's just that human personality is even more complex.
The evolutionary perspective of personality probably makes most sense when considered in conjunction with other perspectives. Evolutionary psychology can seen, for example, as a theoretical platform which underlies the human personality. At birth, everyone starts from scratch, with a unique genotype, some inbuilt instincts (including a temperament), and a pre-wired capacity to learn certain kinds of behaviors. Biological processes, psychodynamics proceses, behavioral processes, social shaping processes, etc. then unfold, interacting with the individual's genotype, to dynamically create the unique psychological characteristics of the individual. All the time, however, this shaping occurs within certain parameters layed down by the genotype, which itself is a synthesized expression of the knowledge of human evolution about what seems to be adaptive, stored and conveyed through genetic code.
The evolutionary perspective is closely related to all other perspectives of personality. Freud, for example, was ultimately famous for being the father of psychology, by revealing that human behavior was driven by unconscious, instinctual forces. Freud understood personality as arising from the way in which humans were able to resolve these instinctual impulses (such as for pleasure, sex, food, etc.) with societal constraints, and the long-term needs of the individual. Indeed, there is evidence that Freud was influenced by the writings of Darwin and that he greatly admired Darwin's work (Sulloway, 1979, cited in McAdams, 1994).
Skinner, the famous advocate for behavioral understandings of the human behavior, also understood their to be a role played by evolutionary forces, although he saw this as not being as important as environmental reinforcement:
"We can trace a small part of human behavior...to natural selection and the evolution of the species, but the greater part of human behavior must be traced to the contingencies of reinforcement, especially to the very complex social contingencies we call cultures. Only when we take those histories into account can we explain why people behave the way they do."
- Skinner, 1989, p. 18, cited in Feist & Feist (2002, p. 281)
Rather than see the various perspectives in opposition from one another, as Skinner seems to do here, I think it is more useful and productive to understand how they all work together in various ways to create the multi-faceted reality of human personality.
The evolutionary perspective, then, views personality as the product of a long history during which it was advantageous for humans to adopt particular characteristic ways of thinking and behaving. Evolutionary forces are most useful for understanding some of the broad trends in apparently instinctual drives. It is also seems that although we have been shaped as a species by the challenge of survival, understanding individual's personalities is often best approached from other perspectives, particularly because the evolutionary perspective currently seems to offer little in the way of practical intervention or assistance in dealing with personality problems.
Variation in human characteristics as adaptive for the species
An important principle of natural selection is that a species will exhibit variations in various physical and behavioral characteristics. In this way, over time, individuals with physical and behavioral characteristics which are most adaptive for survival will be more likely to survive and pass on their characteristics to their off-spring. Over a long period of time, this leads to eventually to entirely different species, or the gradual shaping (evolution) of a species to have some characteristics and not others.
In this light, then, observations of the wide variations in human personality can be understood as the process of evolution throwing up variations of the human psyche which allows the most adaptive personalities to survive more often and procreate.
In a complex species, such as humans, it is also important to realize that quite different personalities may prove adaptive in different ways. For example, highly aggressive behaviors can be adapative in that they allow a person to stand up for themselves and fight for their share, or more, of available resources. However, this also makes a person vulnerable to the aggression of others. So, it is also understandable that more submissive or passive personalities can be adaptive. By avoiding conflict with others, the individual is less likely to suffer direct harm from the aggression of others, but may find that it is difficult to get access to the resources for survival.
For some aspects of personality, there appear to be convincing evolutionary explanations; for other aspects, for other aspects of personality, evolutionary perspectives are less useful. Evolutionary perspectives are probably most useful for explaining general societal behavior trends.
An example of such a trend is that males are greater perpetrators of violence than women.
During human history, it seems males evolved with particular tendencies and capacities that were advantageous for hunting and physical defence of tribes. This underlying predisposition of males seems to also predispose males to also being more likely to have overly violent behaviors. This may be due to higher than normal levels, for example, of particular hormones and neurochemicals (testosterone, for example). Other behavioral sex differences which have attracted evolutionary explanations include the higher rates of promiscuity for males, and the higher rates of rape by males.
Behavioral genetics
Behavioural genetics studies the way inherited biological material i.e. genes, can influence patterns of behaviour.
Behavioural genetics has sometimes been called �trait� genetics as it examines the way our genes influence our personality traits.
The basic methodology of behavioural genetics is to compare similarities in personality between individuals who are and are not genetically related, or who are related to different degrees.
Humans are highly similar to each other genetically. About 90% of human genes are identical from one individual to another. Behavioural genetics concentrates on the approximately 10% of the human genome that does vary.
Behavioural genetics, like trait psychology, focuses exclusively on aspects of personality that differ from one individual to another. The inheritance of species-specific traits or traits that all humans share is examined later in evolutionary psychology.
The basic assumption of behavioural genetics is that if a trait is influenced by genes then it ought to be more highly correlated across pairs of identical (monozygotic:MZ) twins than across pairs of fraternal (dyzygotic

Z) twins, and more highly correlated across closer genetic relatives than across more distant genetic relatives.
Across many personality traits the average correlation across MZ twins is .50 and across DZ twins is .30 (e.g. Bouchard & McGue, 1990; Loehlin & Nichols, 1976). Thus according to twin studies average heritability of most personality traits is .40. This is interpreted to mean that the proportion of behavioural variance that can be explained by genetic variance is 40%. (This is a heritability coefficient i.e. a percentage not a correlation coefficient).
Heritability of the Big Five personality factors
Are the Big 5 traits influenced by our genes? A number of twin studies have explicitly examined the heritability of C, A, O, N, and E.
Evidence of heritability for conscientiousness(Jang, McCrae, Angleimer, Riemann, & Livesey, 1998)
Evidence of heritability for agreeableness (Jang, Livesey, & Vernon, 1996)
Evidence of heritability for openness to experience (Loehlin, 1992)
Strong and consistent evidence of heritability for neuroticism
Strong and consistent evidence of heritability for extraversion